44===new zealand pteridophytes

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New Zealand Journal of Botany, 1985, Vol. 23 : 431-489 OO28-825X/85/23O3-O431$2.5O/O © Crown copyright 1985 431 A revised classification of New Zealand pteridophytes with a synonymic checklist of species P. J. BROWNSEY National Museum, Private Bag Wellington, New Zealand D. R. GIVEN Botany Division, DSIR Private Bag, Christchurch, New Zealand J. D. LOVIS Botany Department, University of Canterbury Christchurch, New Zealand Abstract A revised classification and checklist of New Zealand pteridop
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N e w Z e a la n d J o u r n a l o fB o ta n y , 1 9 8 5 , V o l. 2 3: 4 3 1 -4 8 9 O O 2 8 -8 2 5 X /8 5 /2 3 O 3 -O 4 3 1 $ 2 .5 O /O \u00a9 Crown copyright 1985 4 3 1 A re v is e d c la s s ific a tio n of New Z e a la n d p te rid o p h y te s w ith a s y n o n y m ic c h e c k lis t o f s p e c ie s P . J. B R O W N S E Y N a tio n a l M u s e u m , P riv a te B ag W e llin g to n , N e w Z e a la n d D .R . G IV E N B o ta n y D iv is io n , D S IR P rivate Bag, C hristchurch, N ew Z e a la n d J. D . L O V IS B o ta n y D e p a rtm e n t, U n iv e rs ity o f C a n te rb u ry Christchurch, N ew Z e a la n d A b stract A re v is e d c la s s ific a tio n a n d c h e c k lis t of N ew Zealand pteridophytes is presented, based o n re c e n tly p u b lis h e d p h y le tic schem es fo r th e P te rid o p h y ta , a n d o n re v is io n s o f in d iv id u a l fam ilies and genera. The list com prises 211 species o f w h ic h 2 2 a re a d v e n tiv e a n d 1 8 9 n a tiv e ; th e la tte r include 8 unnam ed species (either undescribed or u n id e n tifie d ) a n d a fu rth e r 6 w h ic h a re s u b d iv id e d into separate subspecies. O f the native taxa, 89 spe- cies (47% ) and 3 subspecies (50% ) are believed to be endem ic to the New Z e a la n d b o ta n ic a l re g io n . Individual taxa are discussed w ith particular refer- ence to recent revisions and areas requiring further s tu d y . S y n o n y m ic a n d a lp h a b e tic a l lis ts a re p ro - v id e d o f a ll v a lid ly p u b lis h e d p te rid o p h y te n a m e s based on New Z e a la n d types or used by authors w ith reference to the N ew Zealand flora. The m ore im p o rta n t s p e c ie s e rro n e o u s ly o r d u b io u s ly recorded for N ew Z e a la n d a re a ls o lis te d . K eyw ords c h e c k lis t; c la s s ific a tio n ; e rro n e o u s records; ferns; N ew Z e a la n d flo ra ; n o m e n c la tu re ; p la n t ta x o n o m y ; p te rid o p h y te s ; s y n o n y m ic lis t IN T R O D U C T IO N The currently accepted standard reference w ork on the New Z e a la n d p te rid o p h y te s is s till th e tre a t- m e n t b y A lla n (1 9 6 1 ) in V o lu m e I o f th e F lo r a o f N ew Zealand . This has since been supplem ented b y tw o w ell-illu strated a n d w id e ly -re a d p u b lic a - tio n s (C ro o k e s 1 9 6 3 , H e a th & C h in n o c k 1 9 7 4 ), b u t R eceived 5O c to b e r1 9 8 4 ; a c c e p te d 29 January 1985 re g re tta b ly n e ith e r o f th e m is c o m p re h e n s iv e in its c o v e ra g e , th e fo rm e r d e a lin g o n ly w ith th e F ilic o p - s id a a n d th e la tte r o m ittin g m a n y o f th e ra re r a n d South Island species. A lla n s w o rk o n th e p te rid o p h y te s , th o u g h n o t p u b lis h e d u n til1 9 6 1 , w a s a c tu a lly c o m p le te d m a n y years previously and w as based on the classifica- tio n p ro p o s e d b y H o lttu m (1 9 4 9 ). H o lttu m him self (1973a) has now revised m any of his earlier ideas w hilst in recent years several other w orkers (N ayar 1 9 7 0 , C rabbe et al. 1975, Lovis 1977, Pichi Ser- m olli 1977) have also produced phyletic schem es o f th e ir o w n w h ic h d e p a r t ra d ic a lly fro m tra d i- tio n a l id e a s o f fe rn e v o lu tio n . In th e lig h t o f th e s e c h a n g in g c o n c e p ts o f fe rn c la s s ific a tio n it is c le a rly d e s ira b le th a t so m e a tte m p t b e m a d e to p ro v id e a m ore m odern arrangem ent for the New Z e a la n d p te rid o p h y te flo ra . O u r c o n c e p ts of fam ilies a n d g e n e ra a re a ls o c h a n g in g a n d in the twenty or m ore years since A lla n s w o rk th e re h a v e b e e n m a jo r re v is io n s o f s u c h fa m ilie s a s H y m e n o p h y lla c e a e (M o rto n 1 9 6 8 ), C yatheaceae (T ry o n 1 9 7 0 ), a n d T h e ly p te rid a c e a e (H o lttu m 1971) as w ell as generic revisions ofL a s - tr e o p s is(T in d a le 1 9 6 5 ), D o o d ia (P a rtis 1 9 7 2 ), T m e- s ip te r is (C h in n o c k 1 9 7 5 ), G ra m m itis (P a rris & G iv e n 1976), L in d s a e a (K ram er & Tindale 1976), A s p le n iu m (B ro w n sey 1977a, b),D eparia (K ato 1 9 8 4 ), a n dH y p o le p is (B ro w n se y & C h in n o c k 1 9 8 4 ). T h e s e , to g e th e r w ith o th e r s m a lle r re v is io n s , a d d i- tio n s o f n e w species, and s u n d ry n o m e n c la tu ra l c h a n g e s , h a v e re s u lte d in a s itu a tio n w h e re a h ig h proportion of the nam es used by A llan (1961) are o u t o f d a te . In recent years, too, New Zealand botanists have b e c o m e in c re a s in g ly aw are o f th e im p o rta n c e o f a d v e n tiv e species. Though th e n u m b e r o f in tro - d u c e d p te rid o p h y te s is s m a ll, th e re a re s o m e w h ic h a re o f sig n ific a n t d is trib u tio n o r e c o n o m ic im p o r- ta n c e . N o n e a re in c lu d e d in A lla n s F lo ra , w h e re o n ly th e n a tiv e s p e c ie s a re d e a lt w ith , b u t th e y h a v e re c e n tly b e e n lis te d b y B ro w n se y (1 9 8 0 ). N o com prehensive s tu d y o f th e N ew Z e a la n d p te rid o p h y te flo ra sim ilar to th o s e p ro d u c e d b y B ro w n lie (1 9 6 9 , 1 9 7 7 ) fo r N e w C a le d o n ia a n d F iji has yet been attem pted, and over tw o decades have e la p s e d s in c e th e p u b lic a tio n o f V o lu m e1 o f A lla n s F lo ra . T h e p u rp o s e o f th e p re s e n t p u b lic a tio n i s th e re fo re t o p ro v id e a n in te rim re v ie w o f th e increased knowledge of New Zealand pteridophyte4 3 2 N ew Zealand Journal of B otany, 1985, V ol. 23 taxonom y and to suggest areas w here further w ork is d e s ira b le . S p e c ific a lly th is in c lu d e s : i) A re v is e d c la s s ific a tio n a n d c h e c k lis t o f N e w Z e a la n d p te rid o p h y te s b a s e d o n o th e r re c e n tly p u b lis h e d p h y le tic s c h e m e s fo r th e P te rid o p h y ta . ii) C o m m e n t o n in d iv id u a l ta x a w ith p a rtic u la r re fe re n c e t o recent revisions and a re a s re q u irin g fu rth e r s tu d y . iii) A s y n o n y m ic lis t o f a ll th e p u b lis h e d p te r id o - p h y te n a m e s e ith e r b a s e d o n N e w Z e a la n d m a te ria l o r u s e d b y a u th o rs w ith re fe re n c e t o th e N ew Zealand flora. iv) A lis t of the m o re im p o rta n t s p e c ie s e rro - neously or dubiously recorded for N ew Z e a la n d . C L A S S IF IC A T IO N OF THE NEW Z E A L A N D P T E R ID O P H Y T A In the last 15 years, at least five phylogenetic class- ific a tio n s o f th e P te r id o p h y ta o r F ilic o p s id a h a v e b e e n p ro p o s e d (N ayar 1970, Crabbe et al. 1975, L o v is 1 9 7 7 , P ic h i S e rm o lli 1 9 7 7 , T ry o n & T ry o n 1 9 8 2 ).D espite m any d iffe re n c e s i n th e d e ta ile d arrangem ents o f fam ilies a n d g e n e ra in th e s e schem es and in the levels of the hierarchical groups adopted, there are som e encouraging signs of broad a g re e m e n t. W ith in the m ore advanced ferns, for e x a m p le , th e c la s s ific a tio n o f w h ic h h a s a lw a y s p ro v e d th e m o s t c o n te n tio u s , th re e m a jo r lin e s o f e v o lu tio n h a v e b e e n re c o g n is e d in a ll b u t th e la s t of the proposed schem es: i) S c h iz a e o id fe rn s le a d in g to th e a d ia n to id f e r n s , ii) G le ic h e n io id fe rn s le a d in g t o th e d ip te ro id , p o ly p o d io id , a n d g ra m m itio id g ro u p s , iii) C y a th e o id , th e ly p te ro id , a n d d e n n s ta e d tio id fe rn s le a d in g to th e a s p le n io id , d ry o p te ro id , d a v - a llio id , a n d b le c h n o id g ro u p s . H o w e v e r, J a r r e tt (1 9 8 0 ) h a s re c e n tly c ritic is e d th e a s s o c ia tio n o f th e P o ly p o d ia c e a e a n d G ra m m iti- d a c e a e w ith th e g le ic h e n io id fe rn s , a n d b e lie v e s th a t they are allied to advanced indusiate fam ilies (see p . 4 3 9 ). H e r id e a s o n th e re la tio n s h ip s o f th e p o ly - podioid ferns are reflected in the essentially con- servative scheme of Tryon & Tryon. P la c e m e n t o f th e film y fe rn s a n d th e ir s a te llite g ro u p s re m a in s th e m a jo r a re a of disagreement betw een the other four schem es, N ayar and Pichi S e rm o lli in c lu d in g th e m in th e c y a th e o id lin e , w h ils t C rabbe e t a l. a n d L o v is re ta in th e m as a q u ite separate group. Of lesser concern are the M arsi- le a le s w h ic h a r e p la c e d a s a n o ff-sh o o t o f th e s c h i- z a e o id lin e b y L o v is a n d P ic h i S e rm o lli b u t le ft undecided by Crabbe et al. and ignored by Nayar. O n ly C rabbe e t a l. a n d P ic h i S erm o lli h a v e tre a te d th e fe rn a llie s in d e ta il b u t th e re a p p e a rs to be no m ajor disagreem ent betw een them \u2014 n e ith e r a u th o r c h o o s in g to in c lu d e th e P s ilo ta le s w ith in th e Filicopsida as suggested by B ierhorst(1 9 7 1 , 1977). Pichi Serm olli has, how ever, rearranged th e fe rn a llie s , m o v in g th e P s ilo to p s id lin e c lo s e r to th e tru e f e r n s . The classification o u tlin e d h e re o w e s s o m e th in g to a ll th e re c e n tly p ro p o s e d schem es but is iden- tic a l to n o n e o f th e m . It is p ro b a b ly c lo s e s t in o v e r- a ll c o n c e p t to th a t o f P ic h i S e rm o lli, b u t d iffe rs in th e tra n s p o s itio n o f th e s c h iz a e o id a n d g le ic h e n io id lin e s ,a n d m ost s ig n ific a n tly in th e h ie ra rc h ic a l le v e ls a d o p te d fo r d iffe re n t ta x a . P ic h i S e rm o lli is m o re d iv is iv e in h is ta x o n o m ic a p p ro a c h th a n u s , w ith the result that his subclasses correspond t o o u r orders, and h is o rd e rs a p p ro x im a te t o o u r fam ilies. In th e circu m scrip tio n of fam ilies (th o u g h not in system atic arrangem ent) our schem e is iden- tic a l w ith th a t o f T ry o n & T ry o n (1 9 8 2 ) e x c e p t fo r th e G ra m m itid a c e a e w h ic h th e y in c lu d e w ith P o ly p o d ia c e a e . N o consideration has been given to subclasses or subfam ilies in this classification, the purpose being sim ply to provide a m ore m odern arrangem ent of th e N e w Z e a la n d ta x a r a th e r th a n a c o m p le te p h y - lo g e n e tic break-dow n. T he la tte r c a n o n ly b e a c h ie v e d b y c o n s id e ra tio n o f a ll th e k n o w n fe rn taxa and is outside the scope of the present paper. G enera a n d s p e c ie s a re a rra n g e d a lp h a b e tic a lly w ith in fam ilies. The list of species accepted here and the choice o f n a m e s fo r th e m are in som e instances purely a m atter o f p e rs o n a l o p in io n b a s e d o n a v a ila b le know ledge at this tim e. Som e com prom ise has been necessary, even betw een the three authors of this paper \u2014 th e q u e s tio n o fC y a th e a v e rs u s A ls o p h ila andS p h a e r o p te r is being perhaps the best exam ple. It is h o p e d , h o w e v e r, th a t th e lis t w ill p ro v id e a re fe re n c e p o in t fo r fu tu re w o rk o n th e N e w Z e a la n d flo ra , a n d th a t th e a c c o m p a n y in g n o te s w ill in d i- c a te w h e re th e R u le s o f N o m e n c la tu re d ic ta te th e a d o p tio n of a p a rtic u la r name, or a lte rn a tiv e ly w h e re a v a lid c h o ic e e x is ts . C H E C K L IS T O F N E W Z E A L A N D P T E R ID O P H Y T A A = species adventive in N ew Z e a la n d C = species confined to the C hatham Is la n d s E = s p e c ie s e n d e m ic to th e N e w Z e a la n d b o ta n ic a l re g io n K = species confined t o th e K erm adec Is la n d s w ith in th e N e w Z e a la n d re g io n P K = s p e c ie s c o n fin e d to th e P o o r K n ig h ts Is la n d sN e w Z e a la n d J o u r n a l o fB o ta n y , 1 9 8 5 , V o l. 2 3: 4 3 1 -4 8 9 O O 2 8 -8 2 5 X /8 5 /2 3 O 3 -O 4 3 1 $ 2 .5 O /O \u00a9 Crown copyright 1985 4 3 1 A re v is e d c la s s ific a tio n of New Z e a la n d p te rid o p h y te s w ith a s y n o n y m ic c h e c k lis t o f s p e c ie s P . J. B R O W N S E Y N a tio n a l M u s e u m , P riv a te B ag W e llin g to n , N e w Z e a la n d D .R . G IV E N B o ta n y D iv is io n , D S IR P rivate Bag, C hristchurch, N ew Z e a la n d J. D . L O V IS B o ta n y D e p a rtm e n t, U n iv e rs ity o f C a n te rb u ry Christchurch, N ew Z e a la n d A b stract A re v is e d c la s s ific a tio n a n d c h e c k lis t of N ew Zealand pteridophytes is presented, based o n re c e n tly p u b lis h e d p h y le tic schem es fo r th e P te rid o p h y ta , a n d o n re v is io n s o f in d iv id u a l fam ilies and genera. The list com prises 211 species o f w h ic h 2 2 a re a d v e n tiv e a n d 1 8 9 n a tiv e ; th e la tte r include 8 unnam ed species (either undescribed or u n id e n tifie d ) a n d a fu rth e r 6 w h ic h a re s u b d iv id e d into separate subspecies. O f the native taxa, 89 spe- cies (47% ) and 3 subspecies (50% ) are believed to be endem ic to the New Z e a la n d b o ta n ic a l re g io n . Individual taxa are discussed w ith particular refer- ence to recent revisions and areas requiring further s tu d y . S y n o n y m ic a n d a lp h a b e tic a l lis ts a re p ro - v id e d o f a ll v a lid ly p u b lis h e d p te rid o p h y te n a m e s based on New Z e a la n d types or used by authors w ith reference to the N ew Zealand flora. The m ore im p o rta n t s p e c ie s e rro n e o u s ly o r d u b io u s ly recorded for N ew Z e a la n d a re a ls o lis te d . K eyw ords c h e c k lis t; c la s s ific a tio n ; e rro n e o u s records; ferns; N ew Z e a la n d flo ra ; n o m e n c la tu re ; p la n t ta x o n o m y ; p te rid o p h y te s ; s y n o n y m ic lis t IN T R O D U C T IO N The currently accepted standard reference w ork on the New Z e a la n d p te rid o p h y te s is s till th e tre a t- m e n t b y A lla n (1 9 6 1 ) in V o lu m e I o f th e F lo r a o f N ew Zealand . This has since been supplem ented b y tw o w ell-illu strated a n d w id e ly -re a d p u b lic a - tio n s (C ro o k e s 1 9 6 3 , H e a th & C h in n o c k 1 9 7 4 ), b u t R eceived 5O c to b e r1 9 8 4 ; a c c e p te d 29 January 1985 re g re tta b ly n e ith e r o f th e m is c o m p re h e n s iv e in its c o v e ra g e , th e fo rm e r d e a lin g o n ly w ith th e F ilic o p - s id a a n d th e la tte r o m ittin g m a n y o f th e ra re r a n d South Island species. A lla n s w o rk o n th e p te rid o p h y te s , th o u g h n o t p u b lis h e d u n til1 9 6 1 , w a s a c tu a lly c o m p le te d m a n y years previously and w as based on the classifica- tio n p ro p o s e d b y H o lttu m (1 9 4 9 ). H o lttu m him self (1973a) has now revised m any of his earlier ideas w hilst in recent years several other w orkers (N ayar 1 9 7 0 , C rabbe et al. 1975, Lovis 1977, Pichi Ser- m olli 1977) have also produced phyletic schem es o f th e ir o w n w h ic h d e p a r t ra d ic a lly fro m tra d i- tio n a l id e a s o f fe rn e v o lu tio n . In th e lig h t o f th e s e c h a n g in g c o n c e p ts o f fe rn c la s s ific a tio n it is c le a rly d e s ira b le th a t so m e a tte m p t b e m a d e to p ro v id e a m ore m odern arrangem ent for the New Z e a la n d p te rid o p h y te flo ra . O u r c o n c e p ts of fam ilies a n d g e n e ra a re a ls o c h a n g in g a n d in the twenty or m ore years since A lla n s w o rk th e re h a v e b e e n m a jo r re v is io n s o f s u c h fa m ilie s a s H y m e n o p h y lla c e a e (M o rto n 1 9 6 8 ), C yatheaceae (T ry o n 1 9 7 0 ), a n d T h e ly p te rid a c e a e (H o lttu m 1971) as w ell as generic revisions ofL a s - tr e o p s is(T in d a le 1 9 6 5 ), D o o d ia (P a rtis 1 9 7 2 ), T m e- s ip te r is (C h in n o c k 1 9 7 5 ), G ra m m itis (P a rris & G iv e n 1976), L in d s a e a (K ram er & Tindale 1976), A s p le n iu m (B ro w n sey 1977a, b),D eparia (K ato 1 9 8 4 ), a n dH y p o le p is (B ro w n se y & C h in n o c k 1 9 8 4 ). T h e s e , to g e th e r w ith o th e r s m a lle r re v is io n s , a d d i- tio n s o f n e w species, and s u n d ry n o m e n c la tu ra l c h a n g e s , h a v e re s u lte d in a s itu a tio n w h e re a h ig h proportion of the nam es used by A llan (1961) are o u t o f d a te . In recent years, too, New Zealand botanists have b e c o m e in c re a s in g ly aw are o f th e im p o rta n c e o f a d v e n tiv e species. Though th e n u m b e r o f in tro - d u c e d p te rid o p h y te s is s m a ll, th e re a re s o m e w h ic h a re o f sig n ific a n t d is trib u tio n o r e c o n o m ic im p o r- ta n c e . N o n e a re in c lu d e d in A lla n s F lo ra , w h e re o n ly th e n a tiv e s p e c ie s a re d e a lt w ith , b u t th e y h a v e re c e n tly b e e n lis te d b y B ro w n se y (1 9 8 0 ). N o com prehensive s tu d y o f th e N ew Z e a la n d p te rid o p h y te flo ra sim ilar to th o s e p ro d u c e d b y B ro w n lie (1 9 6 9 , 1 9 7 7 ) fo r N e w C a le d o n ia a n d F iji has yet been attem pted, and over tw o decades have e la p s e d s in c e th e p u b lic a tio n o f V o lu m e1 o f A lla n s F lo ra . T h e p u rp o s e o f th e p re s e n t p u b lic a tio n i s th e re fo re t o p ro v id e a n in te rim re v ie w o f th e increased knowledge of New Zealand pteridophyteB row nsey et al.— C lassification of N ew Z e a la n d p te rid o p h y te s 4 3 3 P S IL O T O P S ID A P S IL O T A L E S P s ilo ta c e a e P s ilo tu m n u d u m (L .) P . B e a u v . T m e s ip te r is e lo n g a ta P. A . D angeard la n c e o la ta P. A . D angeard s ig m a tifo liaC h in n o c k ta n n e n s is (Sprengel) Bernh. L Y C O P S ID A L Y C O P O D IA L E S L y c o p o d ia c e a e L y c o p o d iu m a u s tr a lia n u m H erter c e r n u u m L . d e u te r o d e n s u m H erter fa s tig ia tu m R . B r. la te r a le R . B r. ram ulosum K irk s c a r io s u m F o r s t . f. s e r p e n tin u m K unze v a r iu m R . B r. v o lu b ileF o r s t . f. P h y llo g lo s s u m d r u m m o n d iiK unze (E ) (E ) S E L A G IN E L L A L E S S e la g in e lla c e a e S e la g in e lla k r a u s s ia n a(K unze) A . B raun m o e lle n d o r ffii H ie ro n . sp . (W hangarei Falls) IS O E T A L E S Is o e ta c e a e h o e te s a lp in u sK irk k ir k iiA . B raun E Q U IS E T O P S ID A E Q U IS E T A L E S E q u is e ta c e a e E q u is e tu m a r v e n s eL . (A ) (A ) (A ) (E ) (E ) (A ) F IL IC O P S ID A O P H IO G L O S S A L E S O phioglossaceae B o tr y c h iu m a u s tr a leR . B r. biform eC o l. (E ) lu n a r ia (L .) Sw artz O p h io g lo s s u m c o r ia c e u m A. C unn. p e tio la tu m H ook. M A R A T T IA L E S M a ra ttia c e a e M a r a ttia s a lic in a J. E . S m ith OSM UNDALES O sm undaceae L e p to p te r is h y m e n o p h y llo id e s (A . R ich.) C . Presl (E ) s u p e r b a (C ol.) C . P resl (E ) O s m u n d a r e g a lisL . (A ) Todea barbara (L .) T . M o o re S C H IZ A E A L E S S c h iz a e a c e a e L y g o d iu m a r tic u la tu m A . R ic h . (E ) S c h iz a e a a u s tr a lisG a u d ic h . b ifid a W illd . d ic h o to m a(L .) J . E . S m ith fis tu lo s aL a b ill. P te rid a c e a e A d ia n tu m a e th io p ic u mL . c a p illu s -v e n e r isL . (A ) c u n n in g h a m iiH ook. (E ) d ia p h a n u m B lu m e form osum R . B r. fu lv u m R aoul (E ) h is p id u lu m S w artz r a d d ia n u m C . Presl (A ) v ir id e s c e n sC o l. (E ) A nogram m a le p to p h y lla(L .) L in k C h e ila n th e s d is ta n s (R . B r.) M ett. s ie b e r iK unze P e lla e afa lc a ta (R . B r.) Fee r o tu n d ifo lia(F o rst. f.) H o o k . (E ) sp . (dry rocky areas) P te r is c o m a n sF o r s t . f. c r e tic aL . (A ) m a c ile n taA . R ic h . (E ) s a x a tilisC arse (E ) trem u laR . B r. G L E IC H E N IA L E S G le ic h e n ia c e a e D ic r a n o p te r is lin e a r is(B u rm . f.) L . U n d e rw . G le ic h e n ia d ic a r p aR . B r. m ic r o p h y llaR . B r. S tic h e r u s c u n n in g h a m ii(H o o k .) C h in g (E ) fla b e lla tu s (R. Br.) H . St. John G ra m m itid a c e a e A n a r th w p te r is la n c e o la ta(H o o k , f.) P ic h i S e rm .(E ) C te n o p te r is h e te w p h y lla(L a b ill.) T in d a le G ra m m itisb illa r d ie r i W illd . c ilia taC o l. (E ) g iv e n ii P a rris (E ) m a g e lla n ic a D esv. subsp. m a g e lla n ic a m a g e lla n ic a D esv. subsp. n o th o fa g e ti P a rris p a ta g o n ic a (C. Chr.) Parris p o e p p ig ia n a (M ett.) Pichi Serm . p s e u d o c ilia taP a rris (E ) r a w lin g s ii P a rris (E ) r ig id aH o m b ro n (E ) P o ly p o d ia c e a e P hym atosorusd iv e r s ifo liu s (W illd .) P ich i S erm . n o v a e -z e la n d ia e (B a k e r) P ic h i S e rm . (E ) s c a n d e n s (Forst. f.) Pichi Serm .
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